import re
regex = r"((?<![^_])ins\d+)_|([a-zA-Z]+[0-9]*)\{([^{}]*)\}:r\.([-()?+\d]+)?(?:_([-()?+\d]+))?"
test = ["EWSR1{ENST00000397938}:r.1_1364_FLI1{ENST00000429175}:r.1046_3051",
	"EML4{ENST00000318522}:r.1_929_EML4{ENST00000318522}:r.903+188_903+220_ALK{ENST00000389048}:r.4080_6220",
	"FUS{ENST00000254108}:r.1_(608)_FUS{ENST00000254108}:r.(819)_937_DDIT3{ENST00000547303}:r.76_872",
	"TCF3{ENST00000262965}:r.1_1795_ins27_PBX1{ENST00000420696}:r.454_6636",
	"EML4{ENST00000318522}:r.?_ALK{ENST00000389048}:r.?"]
res = []
for s in test:
	for match in re.finditer(regex, s):
		tmp = []
		for groupNum in range(0, len(match.groups())):
			if match.group(groupNum+1):
				tmp.append(match.group(groupNum+1))
		res.append(tmp)
print(res)

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